Peer-reviewed publications

For a more up-to-date list, please visit my Google Scholar profile.


Rosenberger, N.M., Hemberger, J., Williams, N.M. Extreme heat increases risk of pollen limitation through reductions in pollen production and pollen quality. Accepted Annals of Botany Plants


Hemberger, J., Bernauer, O., Gaines-Day, H., Gratton, C. Landscape-scale floral resource discontinuity decreases bumble bee occurrence and alters community composition. In press Ecological Applications 10.1002/eap.2907 [PDF]

Hemberger, J. and Gratton C. Floral resource discontinuity contributes to spatial mismatch between pollinator supply and pollination demand in a pollinator-dependent agricultural landscapes. In press Landscape Ecology 10.1007/s10980-023-01707-w [PDF]

Williams, N.M. and Hemberger, J.. Climate, pesticides and landcover drive current and predicted declines of the western bumble bee. Proceedings of the National Academy of Sciences. 120, 7 (2023) 10.1073/pnas.2221692120


Hemberger, J., Rosenberger, N.M, Williams, N.M. Experimental heat waves disrupt bumble bee foraging through direct heat effects and reduced nectar production. Functional Ecology. 00, 1-11 (2022) ( [PDF]

Hemberger, J., Witynski, G., Gratton, C. Floral resource continuity boosts bumble bee colony performance relative to variable floral resources. Ecological Entomology. 47:4, 703-712 (2022) 10.1002/een.13154 [PDF]


Mola, J.M., Hemberger, J., Kochanski, J., Richardson, L.L., Pearse, I. The importance of forests in bumble bee biology and conservation. BioScience. 71:12, 1234-1248 (2021) 10.1093/biosci/biab121 [PDF]

Page, M.L., Nicholson, C.C., Brennan, R., Britzman, A., Hemberger, J., Greer, J., Kahl, H., Müller, U, Peng, Y., Rosenberger, N, Stuligross, C., Wang, L., Yang, L., Williams, N.M. A meta-analysis of single-visit pollinator effectiveness. American Journal of Botany. 108:11, 2196-2207 (2021) 10.1002/ajb2.1764

Hemberger, J., Crossley, M., Gratton, C. Historical decrease in agricultural landscape diversity is associated with shifts in bumble bee species occurrence. Ecology Letters. 24:9, 1800-1813 (2021) 10.1111/ele.13786 [PDF]


Hemberger, J., Frappa, A., Witynski, G., Gratton, C. Saved by the pulse? Separating the effects of total and temporal food abundance on the growth and reproduction of bumble bee microcolonies. Basic and Applied Ecology. 45, 1-11 (2020) 10.1016/j.baae.2020.04.004 [PDF]


Hemberger, J. and Gratton C. Floral resource pulse decreases bumble bee foraging trip duration in central Wisconsin agroecosystem. Ecological Entomology. 43, 447–457 (2018). 10.1111/een.12516 [PDF]

In review or submitted

Hemberger, J. and Williams, Predicting insect resource use in complex landscapes over space and time with a native bumble bee species. In revision Ecology

Hemberger, J. and Williams, N.M. Recent, rapid restructuring of North American bumble bee communities is associated with climate warming. In revision Ecology Letters [pre-print here]

Manuscripts in preparation

Hemberger, J., Nootenboom, C., Lonsdorf, E., Williams, N.M. An inventory of potential: quantifying the value of underutilized land for pollinator conservation and pollination service delivery.

Hemberger, J., Williams, N.M. Increase in extreme heat threatens insect visitation to pollinator dependent crops in the California Central Valley.

Public science communication

The Bumble Bees of Wisconsin: an interactive website to learn about the ecology and conservation of Wisconsin’s bumble bee fauna, as well as how to identify them in the field. Includes both an online guide and PDF field guide. 2018.